The Titanotheres (also known as brontotheres) are extinct family of large, rhinoceros-like mammals that were ancestors of the horse, rhinoceros, and tapir. Titanotheres had horn-like structures on their snout; bony knobs protruded from their skull and were covered with thick skin. Males had larger knobs than females. These herbivores ate soft forest vegetation and were up to 8 feet (2.5 m) tall at the shoulder. Titanotheres had a tiny brain, only as big as a fist. They had four hoofed toes on each front foot and three hoofed toes on each rear foot They lived from the early Eocene until the middle Oligocene (from 58-30 million years ago). Some Titanotheres include Brontops &, Brontotherium (both 8 ft tall, from North America), Dolichorhinus (4 ft tall, from North America), Eotitanops (1.5 ft tall, from North America and Asia), and Embolotherium (8 ft tall, from Mongolia).
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